---
title: "1 Edward Custo Quotes"
description: "Edward Custo has 1 quotes on SMS4Smile. Browse their most insightful sayings."
image: "https://sms4smile.com/quotes/i/57e/edward-custo-192817_animal-behavior_a-year-long-study-by-the-university-of-p_5388.jpg"
---

# Edward Custo

 **1** quotes in **4** categories

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##  Quotes by Edward Custo

> [“A year-long study by the University of Pennsylvania, ending in 2009 and published in the journal Applied Animal Behavior Science (Elsevier), showed…](https://sms4smile.com/quotes/edward-custo/a-year-long-study-by-the-university-of-pennsylvania-ending-in-2009/)

Edward Custo

[Animal](https://sms4smile.com/quotes/about/animal/)

[Image](https://sms4smile.com/quotes/i/57e/edward-custo-192817_animal-behavior_a-year-long-study-by-the-university-of-p_5388.jpg)

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